Took the liberty of looking up the word "lees".
From etymonline.com
lees Look up lees at Dictionary.com
late 14c., from O.Fr. lies, plural of lie "sediment," probably from Celtic (cf. O.Ir. lige "a bed, a lying"), from PIE root *legh- "to lie" (see lie (v.2)).
mother of pearl Look up mother of pearl at Dictionary.com
1510, translating M.L. mater perlarum, with the first element perhaps connected in popular imagination with obsolete mother "a thick substance concreting in liquors; the lees or scum concreted" [Johnson], which is from the root of mud.
feces Look up feces at Dictionary.com
also faeces, c.1400, "dregs," from L. faeces "sediment, dregs," pl. of faex (gen. faecis) "grounds, sediment, lees, dregs," of unknown origin. Specific sense of "human excrement" is from 1630s.
dregs Look up dregs at Dictionary.com
c.1300 (implied in surname Dryngedregges), from O.N. dregg "sediment," from P.Gmc. *drag- (cf. O.H.G. trestir, Ger. Trester "grapeskins, husks"), from PIE *dher- (1) "to make muddy." Replaced O.E. cognate dræst, dærst "dregs, lees." Figurative use is from 1530s.
stale (adj.) Look up stale at Dictionary.com
c.1300, "freed from dregs or lees" (of ale, wine, etc.), i.e. "having stood long enough to clear," cognate with M.Du. stel "stale" (of beer), and probably ultimately from P.Gmc. base *sta- "stand," the source of O.E. standan "to stand," Perhaps via O.Fr. estaler "halt," from Frankish *stal- "position" (see stall (n.1)). The meaning "not fresh" is first recorded late 15c. Figurative sense (of immaterial things) is recorded from 1560s.
Of interest, not one of those definitions mention anything about fermentation.